Hedyotis acuminata (Cham. & Schltdl.) Steud. var. alicarpa (Fosberg) O. Deg. & I. Deg., Phytologia 7, 14, or 15. . Flora of the Hawaiian Islands --Basionym
Type Information
Hedyotis acuminata (Cham. & Schltdl.) Steud. f. alicarpa Fosberg, Bernice P. Bishop Mus. Bull. 174: 82 1943.
O`ahu: Ko`olau Range, western ridge of Kaipapau, forest. 11 October 1931. O. Degener and K.K. Park 11666.
Holotype [BISH]. Isotype(s) [BISH!, G, GH, GRO, K, LCU, MICH, MIN, MO!, NY [2, mfc!], S].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


*Verification: None.

Hedyotis acuminata (Cham. & Schltdl.) Steud. , Nomencl. bot., ed. 2, 1: 726. 1840. Flora of the Hawaiian Islands --Basionym
Type Information
Kadua acuminata Cham. & Schltdl. , Linnaea 4: 163 1829.
O`ahu. 1816- 1817. L. C. A. von Chamisso s.n..
Holotype [LE].
=Kadua acuminata

*Verification: None.
---------------------


*Verification: None.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. alanii Fosberg, Brittonia 8: 170. 1956.
Type information:
Holotype [NY]. O. Degener 19606. 15 May 1949. Maui: [West Maui], Waiehu, SE ridge of Mt. Alani, in dark gulch. Isotype(s) [BISH!, US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. alicarpa Fosberg, Bernice P. Bishop Mus. Bull. 174: 82. 1943.
Type information:
Holotype [BISH]. O. Degener and K.K. Park 11666. 11 October 1931. O`ahu: Ko`olau Range, western ridge of Kaipapau, forest. Isotype(s) [BISH!, G, GH, GRO, K, LCU, MICH, MIN, MO!, NY [2, mfc!], S].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. alicarpoides Fosberg, Brittonia 8: 171. 1956.
Type information:
Holotype [BISH]. W. H. Hatheway 456. Maui: West Maui, first (makai) S branch of Makamakaole Stream.1000ft. Isotype(s) [F, MO, NY, US!].

Note: Holotype missing 8/95 (RKS); not found in general collection (6/97).

=Kadua acuminata
Note: Holotype missing 8/95 (RKS); not found in general collection (6/97).

*Verification: R.K. Shannon.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. basipuba Fosberg, Brittonia 20: 290. 1968.
Type information:
Holotype [US]. O. Degener and I. Degener 26885. 7 February 1960. O`ahu: Kawainui Valley, open woods.1500ft. Isotype(s) [A, B, BM, BISH!, G, L, MASS!, NY [mfc!], PTBG].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. brevimucronata Fosberg, Bernice P. Bishop Mus. Bull. 174: 76. 1943.
Type information:
Holotype [B]. W. Hillebrand s.n.. Kaua`i.

Note: Holotype presumably destroyed.

=Kadua acuminata
Note: Holotype presumably destroyed.

*Verification: None.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. brownii Fosberg, Bernice P. Bishop Mus. Bull. 174: 79. 1943.
Type information:
Holotype [BISH]. O. Degener , K.K. Park and C. Potter, & G. T. Shigeura 11672. 3 November 1935. O`ahu: [Ko`olau Range], Waimalu, shrubby slope.1000ft. Isotype(s) [B, BISH! [2], CU, G, GRO, K, L, LCU, MASS!, MICH, MIN, MO!, NY [2, mfc!], S, US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. calycosa Fosberg, Brittonia 20: 289. 1968.
Type information:
Holotype [US]. O. Degener , I. Degener and & W. Fleming 25128. 27 March 1959. Maui: West Maui, Honokawai Ditch Trail, in shade near stream. Isotype(s) [BISH! [2], NY [3, mfc!], US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. compacta Fosberg, Brittonia 8: 169. 1956.
Type information:
Holotype [US]. O. Degener 22200. 2 March 1952. Lana`i: N (mauka) of Lana`i City, shrubby slope. Isotype(s) [BISH!].
=Kadua acuminata

*Verification: R.K. Shannon.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. cylindrica Fosberg, Bernice P. Bishop Mus. Bull. 174: 90. 1943.
Type information:
Holotype [BISH]. O. Degener 11659. 26 August 1926. Hawai`i: mauka of Punalu`u, in rain forest.
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. duripes Fosberg, Bernice P. Bishop Mus. Bull. 174: 84. 1943.
Type information:
Holotype [BISH]. O. Degener 11731. 21 June 1928. Moloka`i: E of Pauhunui Point, open woods overlooking ocean. Isotype(s) [G, MO!, NY [mfc!]].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. eamesii Fosberg, Bernice P. Bishop Mus. Bull. 174: 86. 1943.
Type information:
Holotype [BISH]. J. F. Rock s.n.. s.d.. Lana`i: ridge above Koele.
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. elliptica Fosberg, Bernice P. Bishop Mus. Bull. 174: 79. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 12956. 1 March 1936. O`ahu: Ko`olau Range, Castle (Pig-god) Trail, Kaluanui, wet forest.650m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. forbesii Fosberg, Bernice P. Bishop Mus. Bull. 174: 88. 1943.
Type information:
Holotype [BISH]. C.N. Forbes 1690.M. 20 November 1919. Maui: East Maui, ridge right side Kipahulu.
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. grandis (A. Gray) Fosberg, Bernice P. Bishop Mus. Bull. 174: 90. 1943. Flora of the Hawaiian Islands --Basionym
Type Information
Kadua grandis A. Gray , Proc. Amer. Acad. Arts 4: 318 1859 [Sept].
Hawai`i: s.l.. 1840. Rémy 390.
Lectotype [GH]. Isolectotype(s) [L [?]]. Lectotypified in Fosberg. Bernice P. Bishop Mus. Bull. 174: 90. 1943.

Note: Gray cited no specimens, but Fosberg considered the Remy specimen to be original material; he apparently only saw two specimens that Gray applied this name to, and his citation of Remy specimen is effectively a lectotypification. St. John's lectotypification is superfluous, in addition to which he implied that there are two U.S. Expl. Exped. collections at US when in fact there is only one. Fosberg indicated that the number on the label of the GH sheet was not clear; he later (Brittonia 8: 171, 1956) commented on a Remy 350 at L that he said is probably a duplicate of the lectotype.

=Kadua acuminata
*Verification: R.K. Shannon, W.L. Wagner.
---------------------


*Verification: R.K. Shannon.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. grayana Fosberg, Bernice P. Bishop Mus. Bull. 174: 79. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 13063. 12 April 1936. O`ahu: Wai`anae Mts., Mokule`ia, Pahole (Kukuiula) Gulch, near head of right branch, moist bottom of gulch.550m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. greenwellae Fosberg, Brittonia 8: 170. 1956.
Type information:
Holotype [NY]. O. Degener , W.H. Hatheway and A. Greenwell 21151. 28 November 1950. Maui: West Maui, S side Makamakole Stream, in rich shrubby forest at narrowest rainy part of gulch.1500ft. Isotype(s) [BISH!, NY [mfc!], US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. halawana Fosberg, Bernice P. Bishop Mus. Bull. 174: 76. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 9458. 30 April 1933. O`ahu: Ko`olau Range, Kalauao-Waimalu Ridge, wet forest.530m. Isotype(s) [US! [2]].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. haupuensis Fosberg, Bernice P. Bishop Mus. Bull. 174: 76. 1943.
Type information:
Holotype [BISH]. H. St. John and F.R. Fosberg 13682. 25 December 1933. Kaua`i: Ha`upu (Hoary Head) Mts., Kipu, moist forest.1000ft.
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. heteripes Fosberg, Bernice P. Bishop Mus. Bull. 174: 85. 1943.
Type information:
Holotype [BISH]. G.C. Munro 250. 27 July 1915. Lana`i: Kaiholena.
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. hirtella Fosberg, Bernice P. Bishop Mus. Bull. 174: 80. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 13634. 28 March 1937. O`ahu: Wai`anae Mts., Wai`anaeuka, N ridge of Pu`u Kumakalii, in patch of woods on steep slope.775m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. hirticalyx Fosberg, Bernice P. Bishop Mus. Bull. 174: 89. 1943.
Type information:
Holotype [BISH]. C.N. Forbes 2430.M. 19 May 1920. Maui: West Maui, Olowalu Valley. Isotype(s) [BISH! [2]].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. honopui Fosberg, Brittonia 8: 171. 1956.
Type information:
Holotype [US]. O. Degener and A. Greenwell 21915. 18 February 1952. Hawai`i: Kohala, Honopui Valley, on forested ledges and cliffs along trail leading to Kaukini Cabin.

Note: No material of this collection found at BISH (type and general collections searched 8/97).

=Kadua acuminata
Note: No material of this collection found at BISH (type and general collections searched 8/97).

*Verification: R.K. Shannon.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. iaoensis Fosberg, Bernice P. Bishop Mus. Bull. 174: 87. 1943.
Type information:
Holotype [BISH]. C.N. Forbes 91.M. June 1910. Maui: West Maui, `Iao Valley. Isotype(s) [BISH!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. isae Fosberg, Brittonia 20: 289. 1968.
Type information:
Holotype [US]. O. Degener , I. Degener and W. Cadenhead 27223. 28 November 1960. Kaua`i: Avocado Pear Valley, SW of Pu`uhinahina, in dark, damp woods. Isotype(s) [BISH! [2], NY [mfc!], US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. kaalae (Wawra) Fosberg, Bernice P. Bishop Mus. Bull. 174: 78. 1943. Flora of the Hawaiian Islands --Basionym
Type Information
Kadua kaalae Wawra , Flora 57: 273 1874.
O`ahu: Wai`anae Mts., Ka`ala. 1869- 1870. H. Wawra 2226.
Holotype [W].
=Kadua acuminata

*Verification: None.
---------------------


*Verification: R.K. Shannon.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. koolauensis Fosberg, Bernice P. Bishop Mus. Bull. 174: 78. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 8980. 30 October 1932. O`ahu: Ko`olau Range, upper Manoa Valley, Manoa Cliff Trail, koa forest.500m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. lanaiensis Fosberg, Bernice P. Bishop Mus. Bull. 174: 85. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 12498. 30 November 1935. Lana`i: head of Hukopoe Gulch, moist forest.700m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. liberipes Fosberg, Bernice P. Bishop Mus. Bull. 174: 75. 1943.
Type information:
Holotype [US]. A.S. Hitchcock 15215. s.d.. Kaua`i: Olokele Gulch.425m.
=Kadua acuminata

*Verification: None.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. microliberipes Fosberg, Brittonia 8: 169. 1956.
Type information:
Holotype [?]. W.H. Hatheway , O. Degener and A. Greenwell 454. 28 November 1950. Maui: West Maui, third S branch Makamakaole Stream from ocean.1500ft. Isotype(s) [BISH, F, US!].

Note: None found in BISH type collection (RKS, 8/95), nor in general collection (6/97). Fosberg cited only one collection, but did not designate which institution the type was at. US sheet (published as "Fo") has "isotype" written in Fosberg's hand; should BISH sheet be considered holotype?

=Kadua acuminata
Note: None found in BISH type collection (RKS, 8/95), nor in general collection (6/97). Fosberg cited only one collection, but did not designate which institution the type was at. US sheet (published as "Fo") has "isotype" written in Fosberg's hand; should BISH sheet be considered holotype?

*Verification: R.K. Shannon.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. obovata Fosberg, Bernice P. Bishop Mus. Bull. 174: 88. 1943.
Type information:
Holotype [BISH]. C.N. Forbes 1645. 15 November 1919. Maui: East Maui, ridge left side Kipahulu Valley. Isotype(s) [BISH! [2]].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. ovalicalyx Fosberg, Bernice P. Bishop Mus. Bull. 174: 88. 1943.
Type information:
Holotype [BISH]. O. Degener 2359. 27 August 1927. Maui: [West Maui], near last ditchman's house on way to Mt. `Eke from N, rain forest. Isotype(s) [B, BISH! [2], CU, G, GH, GRO, K, L, LCU, MASS!, MIN, NY [mfc!], US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. ovalifolia (A. Gray) Fosberg, Bernice P. Bishop Mus. Bull. 174: 87. 1943. Flora of the Hawaiian Islands --Basionym
Type Information
Kadua petiolata A. Gray var. ovalifolia A. Gray, Proc. Amer. Acad. Arts 4: 318 1859 [Sept].
Maui: West Maui, s.l.. 1840. U. S. Expl. Exped. s.n..
Holotype [US].

Note: See St. John (Rhodora 87: 581. 1985).

=Kadua acuminata
*Verification: None.
---------------------


*Verification: R.K. Shannon.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. pilicalyx Fosberg, Bernice P. Bishop Mus. Bull. 174: 82. 1943.
Type information:
Holotype [BISH]. O. Degener 11716. 20 May 1928. Moloka`i: small ravine NW of Maunahui, among shrubs. Isotype(s) [B, BISH!, GH, L, MO!, NY [mfc!], S].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. pololuensis Fosberg, Bernice P. Bishop Mus. Bull. 174: 90. 1943.
Type information:
Holotype [BISH]. O. Degener 11658. 11 August 1926. Hawai`i: E slope of Pololu Valley, forest.
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. quadricosta Fosberg, Bernice P. Bishop Mus. Bull. 174: 84. 1943.
Type information:
Holotype [BISH]. O. Degener 2356. 14 October 1927. Moloka`i: near Brown's Ranch, W of Halawa Valley, open woods. Isotype(s) [NY [mfc!]].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. sherffiana Fosberg, Bernice P. Bishop Mus. Bull. 174: 77. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 12998. 22 March 1936. O`ahu: Wai`anae Mts., Waianae`uka, valley S of E ridge of Pu`u Kalena, moist forest.840m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. skottsbergiana Fosberg, Bernice P. Bishop Mus. Bull. 174: 89. 1943.
Type information:
Holotype [BISH]. J. F. Rock 4599. 18 July 1909. Hawai`i: Kohala Mts., Hi`ilawe, Waipi`o Valley. Isotype(s) [BISH! [2]].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. sphaerocarpa Fosberg, Bernice P. Bishop Mus. Bull. 174: 76. 1943.
Type information:
Holotype [BISH]. U. Faurie 383. January 1910. Kaua`i: Hanalei.
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. st.-johnii Fosberg, Bernice P. Bishop Mus. Bull. 174: 85. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 12479. 30 November 1935. Lana`i: ridge below Pu`u Ali`i between Maunalei drainage and Hauola drainage, moist brush-covered ridge.850m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. storeyi Fosberg, Bernice P. Bishop Mus. Bull. 174: 84. 1943.
Type information:
Holotype [BISH]. H. St. John , Dunn and W. B. Storey 13279. 4 July 1933. Moloka`i: Wailau Valley, Kukuinui Ridge.2000ft.
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. subsessilis Fosberg, Bernice P. Bishop Mus. Bull. 174: 87. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 12602. 3 December 1935. Lana`i: Kaiholena Gulch, wet gulch bottom.750m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. toppingii Fosberg, Bernice P. Bishop Mus. Bull. 174: 77. 1943.
Type information:
Holotype [BISH]. O. Degener , K.K. Park and M. Kwon 11663. 14 November 1931. O`ahu: [Ko`olau Range], eastern part of Waialae Ridge, forest. Isotype(s) [B, BISH!, GH, L, MO!, NY [mfc!], S].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. tuberculata Fosberg, Bernice P. Bishop Mus. Bull. 174: 86. 1943.
Type information:
Holotype [BISH]. G.C. Munro 41. August 1913. Lana`i: s.l..
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. turbinata Fosberg, Bernice P. Bishop Mus. Bull. 174: 89. 1943.
Type information:
Holotype [B]. W. Hillebrand s.n.. August 1870. Maui: West Maui, Olualu [Olowalu].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. variabilis Fosberg, Bernice P. Bishop Mus. Bull. 174: 82. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 13386. 23 December 1936. Moloka`i: sloping plateau E of Mapulehu Valley, wet decadent forest.800m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. violetana Fosberg, Bernice P. Bishop Mus. Bull. 174: 81. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 13603. 21 February 1937. O`ahu: Wai`anae Mts., head of Kawaihapai Gulch, brushy ravine.600m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.

Hedyotis acuminata (Cham. & Schltdl.) Steud. f. wailauensis Fosberg, Bernice P. Bishop Mus. Bull. 174: 85. 1943.
Type information:
Holotype [BISH]. F.R. Fosberg 9606a. 3 July 1933. Moloka`i: N coast E Moloka`i, Wailau Valley, steep slope, base of wet pali.200m. Isotype(s) [US!].
=Kadua acuminata

*Verification: R.K. Shannon, W.L. Wagner.