Type Information
Euphorbia celastroides Boiss. var. haupuana Sherff, Bot. Gaz. (Crawfordsville) 97: 594 1936.
Kaua`i, Haupu Lihu`e. 18 March 1909. Rock 2444.
Holotype [BISH]. Isotype(s) [GH].
Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. humbertii Sherff, Bot. Gaz. (Crawfordsville) 97: 596 1936.
Kaua`i 1851- 1855. Rémy 595.
Holotype [GH]. Isotype(s) [BISH?, GH, P [2 sheets]].

Note: Additional syntype: Remy 595, Ni`ihau. Koutnik made the Ni`ihau collection the lectotype of E. celastroides. Although Boissier did not cite numbers for the Remy collections, it is presumed that they are both number 595. The BISH sheet has neither locality nor number on it, so it cannot be determined whether it is an isolectotype of E. celastroides or an isotype of E. celastroides var. humbertii, if either (RKS, 8/95). Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. ingrata O. Deg. & Sherff, in Sherff, Bot. Leafl. 9: 7 1954.
Maui, West Maui, Kahakuloa, grassy windswept promontory. 28 February 1952. Degener 21967.
Holotype [F]. Isotype(s) [BISH!, K, NY [mfc!]].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. kaenana Sherff, Bot. Gaz. (Crawfordsville) 97: 594 1936.
O`ahu, Kaena uplands. 28 March 1915. MacCaughey s.n..
Holotype [BISH].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. kealiana Sherff, Bot. Gaz. (Crawfordsville) 97: 595 1936.
Kaua`i, Ke`alia, in rupibus littoris. Faurie 477.
Holotype [BM]. Isotype(s) [A].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. kohalana O. Deg. & Sherff, in Sherff, Bot. Gaz. (Crawfordsville) 97: 595 1936.
Hawai`i, Kohala. 22 March 1930. Degener 8037.
Holotype [G]. Isotype(s) [BISH! [3], CU, F, LCU, MASS!, MICH, MO!, NY [mfc!]].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. laehiensis O. Deg., I. Deg. & Sherff, in Sherff, Occas. Pap. Bernice P. Bishop Mus. 23(7): 122 1964.
Lana`i, Laehi, only on calcareous, rounded slope. 100 ft. 26 January 1964. Degener and Degener 28760.
Holotype [F]. Isotype(s) [A, B, BISH!, CU, E, F, G, K, L MASS!, MICH, MIN, NY [mfc!], MO!, US!].

Note: Koutnik (1985) erroneously published type as Degener & Degener 28375 (RKS, 8/95). Label on MO sheet indicates that Degener & Degener made the combination in Chamaesyce--check lit. to confirm that it was not published by them. Koutnik, D.L. 1985. Brittonia 37:397-399
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. limahuliensis H. St. John, Phytologia 64: 487 1988.
Kaua`i, Limahuli Valley, SW end, on exposed slopes. 1400 to 1700 ft. 12 April 1978. Perlman 9.
Holotype [BISH]. Isotype(s) [BISH! [5]].

Note: Why so many sheets at BISH? Any dups. anywhere else? R.K. Shannon, D.R. Herbst
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. lorifolia (A. Gray) Sherff, Bot. Gaz. (Crawfordsville) 97: 597 1936.


Note: Note also, this variety publ. ex Skottsb., Acta Horti Gothob. 10: 122. 1936. [D.M. Mix]
*Verification: d.
---------------------

Euphorbia celastroides Boiss. var. mauiensis Sherff, Bot. Gaz. (Crawfordsville) 97: 601 1936.
Maui, sandy isthmus. 1864- 1865. Mann and Brigham 389.
Holotype [US]. Isotype(s) [BISH, CU, F, GH, MO!].

Note: Nom. illeg. This collection was used as the type for Euphorbia multiformis var. lorifolia by Gray. [W.L. Wagner, D.M. Mix]
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. moomomiana Sherff, Bot. Gaz. (Crawfordsville) 97: 593 1936.
Moloka`i, Mo`omomi sands. April 1918. Rock 14014.
Holotype [BISH]. Isotype(s) [BISH! [3], GH, K, US!].

Note: Rock's name Euphorbia munroi was apparently never published. Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. nelsonii H. St. John, Pacific Sci. 30: 15 1976.
1792- 1794. Menzies and Nelson s.n..
Holotype [BM].

*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. nematopoda Sherff, Bot. Gaz. (Crawfordsville) 97: 597 1936.
Kaua`i, Ha`upu Range, left hand side of Kipu Kai Gap. 1 November 1916. Forbes 726.K.
Holotype [BISH].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: d.
---------------------

Euphorbia celastroides Boiss. var. niuensis Sherff, Bot. Gaz. (Crawfordsville) 97: 598 1936.
O`ahu, Niu. Hillebrand s.n..
Holotype [B].

Note: Holotype presumably destroyed; photo F.
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. odonatoides O. Deg. & Sherff, in Sherff, Amer. J. Bot. 38: 55 1951.
Lana`i, Waiakeakua, in decadent forest. 2500 ft. 4 August 1949. Degener and Murashige 19779.
Holotype [F]. Isotype(s) [BISH!, BM, CU, DS, F [photo US!], GH, K, M, MIN, MO!, NY [4, mfc!], US! [2]].

Note: Additional herbaria: CM, G-DEL, LA, P, PH. There is a photo of one of the NY sheets at F. Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. pseudoniuensis O. Deg. & Sherff, in Sherff, Amer. J. Bot. 43: 475 1956.
Kaua`i, Koke`e, Nu`alolo Trail, open ridge. 23 September 1955. Degener et al. 23989.
Holotype [F]. Isotype(s) [BISH! [2], F, K, NY [2, mfc!]].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. saxicola O. Deg. & Sherff, in Sherff, Bot. Gaz. (Crawfordsville) 97:596 1936.
Hawai`i, Kohala, along rocky shore. 9 August 1926. Degener 8088.
Holotype [G]. Isotype(s) [BISH! [2], CU, F, LCU].

*Verification: Koutnik, D.L. 1987. Allertonia 4:331-388.
---------------------

Euphorbia celastroides Boiss. var. stokesii (C. N. Forbes) Sherff, Bot. Gaz. (Crawfordsville) 97: 593 1936.

*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. tomentella (Boiss.) Oudejans, Phytologia 67: 44 1989.

*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. waikoluensis Sherff, Bot. Gaz. (Crawfordsville) 97: 594 1936.
Moloka`i, Kalaupapa and Waikolu along the shore. 26 March 1910. Rock 6191.
Holotype [F]. Isotype(s) [BISH! [3], F, GH].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. hathewayi Sherff, Bot. Leafl. 3:4 1951.
O`ahu, Mokuleia, Wai`anae Mts, east slope of Keawapilau Gulch. 1400 ft. 5 September 1950. Hatheway 223.
Holotype [F]. Isotype(s) [BISH, NY, US!].

*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. , in de Candolle, Prodr. 15(2): 11 1862.
Ni`ihau 1851- 1855. Rémy 595.
Lectotype [P]. Isolectotype(s) [BISH?].

Note: Additional syntype: Remy 595, Kaua`i. Sherff made the Kaua`i collection the type of E. celastroides var. humbertii. Although Boissier did not cite numbers for the Remy collections, it is presumed that they are both number 595. The BISH sheet has neither locality nor number on it, so it cannot be determined whether it is an isolectotype of E. celastroides or an isotype of E. celastroides var. humbertii, if either (RKS, 8/95). Was this lectotypified by Sherff, 1937, Annals of the Missouri Botanical Garden? WLW sez, "Need to see material." Koutnik cites Remy 595 Ni`ihau, but does the number 595 really go with both the Ni`ihau and the Kaua`i collection (Koutnik is the only one who says 595 for the Ni`ihau sheet). Is the publication of var. humbertii (Bot. Gaz. 97: 596. 1936.) using syntype material of var. celastroides 1) effective lectotypification of var. celastroides or 2) invalid publication of var. humbertii?? (11/95). Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: None.
---------------------

Euphorbia celastroides Boiss. f. kahanana Sherff, Bot. Gaz. (Crawfordsville) 97: 596 1936.
O`ahu, W slope of Kahana Valley, sunny ridge. 26 February 1929. Degener and Krauss 8036.
Holotype [BISH]. Isotype(s) [F, NY [mfc!]].

Note: Discrepancies between BISH sheet, Sherff publication, and Koutnik as to where the holotype is. BISH sheet is considered holotype (ex herb. Degener) even though it is labeled "isotype." See Sherff's later citation in Ann. Missouri Bot. Gard. 25: 31 (1938) ("type, Degener; cotypes, Field, N. Y."). R.K. Shannon
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. amplectens Sherff, Bot. Gaz. (Crawfordsville) 97: 598 1936.
Moloka`i, Kalua`uha Valley. August 1912. Forbes 355.Mo.
Holotype [F]. Isotype(s) [BISH! [3], F, K, MIN, MO!, US!].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. arenisaxosa H. St. John, Phytologia 63: 466 1987.
Moloka`i, Kalani, Kaluakoi, sand dunes. 200 ft. 24 December 1948. St. John et al. 23486.
Holotype [BISH]. Isotype(s) [BISH! [2], US!].

Note: Additional collector: J. H. Maysilles.
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. halawana Sherff, Bot. Gaz. (Crawfordsville) 97: 596 1936.
Moloka`i, E end, near Halawa. April 1918. Rock 14041.
Holotype [BISH]. Isotype(s) [BISH! [3], A].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388
*Verification: David Lorence.
---------------------

Euphorbia celastroides Boiss. var. hanapepensis Sherff, Bot. Gaz. (Crawfordsville) 97: 597 1936.
Kaua`i, along the Hanapepe River near the falls. 700 ft. 2-8 July 1895. Heller 2429.
Holotype [MO]. Isotype(s) [US!].

Note: Collection 2429, pro parte. Other collections distributed under Heller 2429 were made 24-26 June (A, GH, BISH! [3], MIN!, NY [mfc!]), and are not primary type material. Sherff clearly distinguished between specimens of Heller 2429 collected 24-26 June and those collected 2-8 July. He cited only the US sheet as an isotype. Possible isotypes: CU, MSC. The paratypes cited by Sherff (F [2], GH, K, NY) probably all represent the same collection, although the locality on the K sheet is given as "Hanapepe and Wahiawa watershed," and the date is 24 June. Koutnik (1987) erroneously considered specimens at BISH, F, GH, K, and MIN to be isotypes. Wagner, W.L. & R.K. Shannon. 1999. Brittonia 51:422-438
*Verification: David Lorence.
---------------------