Smithsonian National Museum of Natural History

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Jun Wen

Jun Wen

Research Botanist & Curator

Phone: 202-633-4881
Fax: 202-786-2563
E-mail Address:

Postal Address
(Mail and Packages):

Smithsonian Institution
PO Box 37012
Washington, DC 20013-7012

Courier Address:
National Museum of Natural History
West Loading Dock
10th and Constitution Ave, NW
Washington, DC 20560-0166

Research Interests
     Monographic, phylogenetic, biogeographic and ethnobotanical studies of the ginseng family (Araliaceae) and the grape family (Vitaceae); Systematics and biogeography of intercontinental disjunct plants in the Northern Hemisphere; Biogeography of Asia; Systematics and conservation of medicinal plants of Asia; Biodiversity inventory and conservation studies in Asia.

Recent Publications - (All Publications)

Duan, Lei, Wen, Jun, Yang, Xue, Liu, Pei-Liang, Arslan, Emine, Ertugrul, Kuddisi and Chang, Zhao-Yang 2015. Phylogeny of Hedysarum and tribe Hedysareae (Leguminosae: Papilionoideae) inferred from sequence data of ITS, matK, trnL-F and psbA-trnH. Taxon, 64(1): 49-64. doi:10.12705/641.26

Ge, Song and Wen, Jun 2015. Journal of Systematics and Evolution: Looking forward. Journal of Systematics and Evolution, 53(1): 1 doi:10.1111/jse.12139

Harris, A. J., Lutz, Sue, Acevedo-Rodríguez, Pedro and Wen, Jun 2015. The utility of the morphological variation of pollen for resolving the evolutionary history of Billia (subfam. Hippocastanoideae, Sapindaceae). Journal of Systematics and Evolution, 53(3): 228-238. doi:10.1111/jse.12130

Huang, Yong-Jiang, Liu, Yu-Sheng, Wen, Jun and Quan, Cheng 2015. First fossil record of Staphylea L. (Staphyleaceae) from North America, and its biogeographic implications. Plant Systematics and Evolution, : 1-16. doi:10.1007/s00606-015-1224-z

Ickert-Bond, Stefanie, Gerrath, Jean M., Posluszny, Usher and Wen, Jun 2015. Inflorescence development in the Vitis–Ampelocissus clade of Vitaceae: the unusual lamellate inflorescence of Pterisanthes. Botanical Journal of the Linnean Society, 179(4): 725-741. doi:10.1111/boj.12348

Kim, Changkyun, Deng, Tao, Wen, Jun, Nie, Ze Long and Sun, Hang 2015. Systematics, biogeography, and character evolution of Deutzia (Hydrangeaceae) inferred from nuclear and chloroplast DNA sequences. Molecular phylogenetics and evolution, 87: 91-104. doi:10.1016/j.ympev.2015.03.002

Liu, Xiu-Qun, Ickert-Bond, Stefanie, Nie, Ze-Long, Zhou, Zhuo, Chen, Long-Qing and Wen, Jun 2015. Phylogeny of the Ampelocissus–Vitis clade in Vitaceae supports the New World origin of the grape genus. Molecular phylogenetics and evolution, doi:10.1016/j.ympev.2015.10.013

Lu, Jin-Mei, Zhang, Ning, Du, Xin-Yu, Wen, Jun and Li, De-Zhu 2015. Chloroplast phylogenomics resolves key relationships in ferns. Journal of Systematics and Evolution, doi:10.1111/jse.12180

Nie, Ze-Long, Funk, Vicki A., Meng, Ying, Deng, Tao, Sun, Hang and Wen, Jun 2015. Recent assembly of the global herbaceous flora: evidence from the paper daisies (Asteraceae: Gnaphalieae). New Phytologist, doi:10.1111/nph.13740

Wang, Hua-Feng, Landrein, Sven, Dong, Wen-Pan, Nie, Ze-Long, Kondo, Katsuhiko, Funamoto, Tsuneo, Wen, Jun and Zhou, Shi-Liang 2015. Molecular phylogeny and biogeographic diversification of linnaeoideae (Caprifoliaceae s. l.) disjunctly distributed in Eurasia, North America and Mexico. PloS One, 10(3): 1-26. doi:10.1371/journal.pone.0116485

Wen, Jun, Ickert-Bond, Stefanie, Appelhans, Marc S., Dorr, Laurence J. and Funk, Vicki A. 2015. Collections-based systematics: opportunities and outlook for 2050. Journal of Systematics and Evolution, doi:10.1111/jse.12181

Wen, Jun, Liu, Jianquan, Ge, Song, Xiang, Qiu-Yun (Jenny) and Zimmer, Elizabeth Anne 2015. Phylogenomic approaches to deciphering the tree of life. Journal of Systematics and Evolution, 53(5): 369-370. doi:10.1111/jse.12175

Xiang, Jian-Ying, Wen, Jun and Peng, Hua 2015. Evolution of the eastern Asian–North American biogeographic disjunctions in ferns and lycophytes. Journal of Systematics and Evolution, 53(1): 2-32. doi:10.1111/jse.12141

Xu, Xin-Wei, Wu, Jin-Wei, Qi, Mei-Xia, Lu, Qi-Xiang, Lee, Peter F., Lutz, Sue, Ge, Song and Wen, Jun 2015. Comparative phylogeography of the wild-rice genus Zizania (Poaceae) in eastern Asia and North America. American Journal of Botany, 102(2): 239-247. doi:10.3732/ajb.1400323

Yi, Ting-shuang, Jin, Gui-hua and Wen, Jun 2015. Chloroplast capture and intra- and inter-continental biogeographic diversification in the Asian – New World disjunct plant genus Osmorhiza (Apiaceae). Molecular phylogenetics and evolution, 85: 10-21. doi:10.1016/j.ympev.2014.09.028

Zhang, Jian-Qiang, Meng, Shi-Yong, Wen, Jun and Rao, Guang-Yuan 2015. DNA Barcoding of Rhodiola (Crassulaceae): a Case Study on a Group of Recently Diversified Medicinal Plants from the Qinghai-Tibetan Plateau. PloS One, 10(3): 1-15. doi:10.1371/journal.pone.0119921

Zhang, Ning, Wen, Jun and Zimmer, Elizabeth Anne 2015. Expression patterns of AP1, FUL, FT and LEAFY orthologs in Vitaceae support the homology of tendrils and inflorescences throughout the grape family. Journal of Systematics and Evolution, 53(5): 469-476. doi:10.1111/jse.12138

Zimmer, Elizabeth Anne and Wen, Jun 2015. Using nuclear gene data for plant phylogenetics: Progress and prospects II. Next-gen approaches. Journal of Systematics and Evolution, 53(5): 371-379. doi:10.1111/jse.12174

Zuo, Yun-Juan, Wen, Jun, Ma, Jin-Shuang and Zhou, Shi-Liang 2015. Evolutionary radiation of the Panax bipinnatifidus species complex (Araliaceae) in the Sino-Himalayan region of eastern Asia as inferred from AFLP analysis. Journal of Systematics and Evolution, 15(3): 210-220. doi:10.1111/jse.12119

Appelhans, Marc S., Wen, Jun and Wagner, Warren L. 2014. A molecular phylogeny of Acronychia, Euodia, Melicope and relatives (Rutaceae) reveals polyphyletic genera and key innovations for species richness. Molecular Phylogenetics and Evolution, 79: 54-68. doi:10.1016/j.ympev.2014.06.014

Appelhans, Marc S., Wen, Jun, Wood, Kenneth R., Allan, Gerard J., Zimmer, Elizabeth Anne and Wagner, Warren L. 2014. Molecular phylogenetic analysis of Hawaiian Rutaceae (Melicope, Platydesma and Zanthoxylum) and their different colonization patterns. Botanical Journal of the Linnean Society, 174(3): 425-448. doi:10.1111/boj.12123

Chin, Siew-Wai, Shaw, Joey, Haberle, Rosemarie, Wen, Jun and Potter, Daniel 2014. Diversification of almonds, peaches, plums and cherries – molecular systematics and biogeographic history of Prunus (Rosaceae). Molecular Phylogenetics and Evolution, 76: 34-48. doi:10.1016/j.ympev.2014.02.024

Ickert-Bond, Stefanie M., Gerrath, Jean and Wen, Jun 2014. Gynoecial Structure of Vitales and Implications for the Evolution of Placentation in the Rosids. International Journal of Plant Sciences, 175(9): 998-1032. doi:10.1086/678086

Li, Rong and Wen, Jun 2014. Phylogeny and biogeography of Asian Schefflera (Araliaceae) based on nuclear and plastid DNA sequences data. Journal of Systematics and Evolution, 52(4): 431-449. doi:10.1111/jse.12052

Liu, Q., Liu, H., Wen, Jun and Peterson, Paul M. 2014. Infrageneric phylogeny and temporal divergence of Sorghum (Andropogoneae, Poaceae) based on low-copy nuclear and plastid sequences. PLoS ONE, 9(8): 1-14. doi:10.1371/journal.pone.0104933

Liu, Qing, Jiang, Bin, Wen, Jun and Peterson, Paul M. 2014. Low-copy nuclear gene and McGISH resolves polyploid history of Eleusine coracana and morphological character evolution in Eleusine. Turkish Journal of Botany, 38: 1-12. doi:10.3906/bot-1305-12

Meng, Ying, Nie, Ze-Long, Deng, Tao, Wen, Jun and Yang, Yong-Ping 2014. Phylogenetics and evolution of phyllotaxy in the Solomon's seal genus Polygonatum (Asparagaceae: Polygonateae). Botanical Journal of the Linnean Society, 176(4): 435-451. doi:10.1111/boj.12218

Triplett, Jimmy K., Clark, Lynn G., Fisher, Amanda E. and Wen, Jun 2014. Independent allopolyploidization events preceded speciation in the temperate and tropical woody bamboos. New Phytologist, 204(1): 66-73. doi:10.1111/nph.12988

Valcárcel, Virginia, Fiz-Palacios, Omar and Wen, Jun 2014. The origin of the early differentiation of Ivies (Hedera L.) and the radiation of the Asian Palmate group (Araliaceae). Molecular Phylogenetics and Evolution, 70: 492-503. doi:10.1016/j.ympev.2013.10.016

Wen, Jun, Boggan, John and Nie, Ze-Long 2014. Synopsis of Nekemias Raf., a segregate genus from Ampelopsis Michx. (Vitaceae) disjunct between eastern/southeastern Asia and eastern North America, with ten new combinations. PhytoKeys, 42: 11-19. doi:10.3897/phytokeys.42.7704

Wen, Jun, Zhang, Jian-Qiang, Nie, Ze-Long, Zhong, Yang and Sun, Hang 2014. Evolutionary diversifications of plants on the Qinghai-Tibetan Plateau. Frontiers in Genetics, 5 doi:10.3389/fgene.2014.00004

Xie, Lei, Yang, Zhi-Yun, Wen, Jun, Li, De-Zhu and Yi, Ting-Shuang 2014. Biogeographic history of Pistacia (Anacardiaceae), emphasizing the evolution of the Madrean-Tethyan and the eastern Asian-Tethyan disjunctions. Molecular Phylogenetics and Evolution, 77: 136-146. doi:10.1016/j.ympev.2014.04.006

Zhang, Jian-Qiang, Meng, Shi-Yong, Allen, Geraldine A., Wen, Jun and Rao, Guang-Yuan 2014. Rapid radiation and dispersal out of the Qinghai-Tibetan Plateau of an alpine plant lineage Rhodiola (Crassulaceae). Molecular Phylogenetics and Evolution, 77: 147-158. doi:10.1016/j.ympev.2014.04.013

Zhang, Jian-Qiang, Meng, Shi-Yong, Wen, Jun and Rao, Guang-Yuan 2014. Phylogenetic Relationships and Character Evolution of Rhodiola (Crassulaceae) based on Nuclear Ribosomal ITS and Plastid trnL-F and psbA-trnH Sequences. Systematic Botany, 39(2): 441-451. doi:10.1600/036364414X680753

Zhou, Shiliang, Dong, Wenpan, Chen, Xiaoqing, Zhang, Xianchun, Wen, Jun and Schneider, Harald 2014. How many species of bracken (Pteridium) are there? Assessing the Chinese brackens using molecular evidence. Taxon, 63(3): 509-521. doi:10.12705/633.9

Chin, Siew-wai, Lutz, Sue M., Wen, Jun and Potter, Daniel 2013. The Bitter and the Sweet: Inference of Homology and Evolution of Leaf Glands in Prunus (Rosaceae) through Anatomy, Micromorphology, and Ancestral–Character State Reconstruction. International Journal of Plant Sciences, 174(1): 27-46. doi:10.1086/668219

Deng, Tao, Zhang, Dai-Gui, Liu, Zhiwei, Tucker, Gordon C., Sun, Hang, Wen, Jun and Nie, Ze-Long 2013. Oxalis wulingensis (Oxalidaceae), an unusual new species from central China. Systematic Botany, 38(1): 154-161. doi:10.1600/036364413X661953

Fan, Deng-Mei, Chen, Jia-Hui, Meng, Ying, Wen, Jun, Huang, Jin-Ling and Yang, Yong-Ping 2013. Molecular phylogeny of Koenigia L. (Polygonaceae: Persicarieae): Implications for classification, character evolution and biogeography. Molecular phylogenetics and evolution, 69(3): 1093-1100. doi:10.1016/j.ympev.2013.08.018

Harris, Aj, Wen, Jun and Xiang, Qiu-Yun 2013. Inferring the biogeographic origins of inter-continental disjunct endemics using a Bayes-DIVA approach. Journal of Systematics and Evolution, 51(2): 117-133. doi:10.1111/jse.12007

Huang, Wei-Ping, Sun, Hang, Deng, Tao, Razafimandimbison, Sylvain G., Nie, Ze-Long and Wen, Jun 2013. Molecular phylogenetics and biogeography of the eastern Asian-eastern North American disjunct Mitchella and its close relative Damnacanthus (Rubiaceae, Mitchelleae). Botanical Journal of the Linnean Society, 171(2): 395-412. doi:10.1111/j.1095-8339.2012.01321.x

Ickert-Bond, Stefanie and Wen, Jun 2013. A taxonomic synopsis of Altingiaceae with nine new combinations. PhytoKeys, 31: 21-61. doi:10.3897/phytokeys.31.6251

Li, Rong, Ma, Peng-Fei, Wen, Jun and Yi, Ting-Shuang 2013. Complete Sequencing of Five Araliaceae Chloroplast Genomes and the Phylogenetic Implications. PLoS ONE, 8(10): e78568 doi:10.1371/journal.pone.0078568

Li, Rong and Wen, Jun 2013. Phylogeny and Biogeography of Dendropanax (Araliaceae), an Amphi-Pacific Disjunct Genus between Tropical/Subtropical Asia and the Neotropics. Systematic Botany, 38(2): 536-551. doi:10.1600/036364413X666606

Liu, Xiao-Lin, Wen, Jun, Nie, Ze-Long, Johnson, Gabriel, Liang, Zong-Suo and Chang, Zhao-Yang 2013. Polyphyly of the Padus group of Prunus (Rosaceae) and the evolution of biogeographic disjunctions between eastern Asia and eastern North America. Journal of Plant Research, 126(3): 351-361. doi:10.1007/s10265-012-0535-1

Lu, Limin, Wang, Wei, Chen, Zhiduan and Wen, Jun 2013. Phylogeny of the non-monophyletic Cayratia Juss. (Vitaceae) and implications for character evolution and biogeography. Molecular phylogenetics and evolution, 68: 502-515. doi:10.1016/j.ympev.2013.04.023

Manchester, Steven R., Kapgate, Dashrath K. and Wen, Jun 2013. Oldest fruits of the grape family (Vitaceae) from the Late Cretaceous Deccan Cherts of India. American Journal of Botany, 100(9): 1849-1859. doi:10.3732/ajb.1300008

Molina, Jeanmaire E., Wen, Jun and Struwe, Lena 2013. Systematics and biogeography of the non-viny grape relative Leea (Vitaceae). Botanical Journal of the Linnean Society, 171(2): 354-376. doi:10.1111/j.1095-8339.2012.01320.x

Nie, Ze-Long, Deng, Tao, Meng, Ying, Sun, Hang and Wen, Jun 2013. Post-Boreotropical dispersals explain the pantropical disjunction in Paederia (Rubiaceae). Annals of Botany, 111(5): 873-886. doi:10.1093/aob/mct053

Ren, Zhumei, Zhong, Yang, Kurosu, Utako, Aoki, Shigeyuki, Ma, Enbo, von Dohlen, Carol D. and Wen, Jun 2013. Historical biogeography of Eastern Asian-Eastern North American disjunct Melaphidina aphids (Hemiptera: Aphididae: Eriosomatinae) on Rhus hosts (Anacardiaceae). Molecular Phylogenetics and Evolution, 69(3): 1146-1158. doi:10.1016/j.ympev.2013.08.003

Shi, Wenting, Wen, Jun and Lutz, Susan 2013. Pollen morphology of the Maddenia clade of Prunus and its taxonomic and phylogenetic implications. Journal of Systematics and Evolution, 51(2): 164-183. doi:10.1111/j.1759-6831.2012.00233.x

Wen, Jun, Kiapranis, Robert and Lovave, Michael 2013. Ampelocissus asekii J. Wen, R. Kiapranis & M. Lovave, a new species of Vitaceae from Papua New Guinea. Phytokeys, 21: 1-6. doi:10.3897/phytokeys.21.4512

Wen, Jun, Lu, L. M. and Boggan, J. K. 2013. Diversity and evolution of Vitaceae in the Philippines. Philippine Journal of Science, 142(Special Issue): 223-244.

Wen, Jun, Ree, Richard H., Ickert-Bond, Stefanie, Nie, Zelong and Funk, Vicki A. 2013. Biogeography: Where do we go from here?. Taxon, 62(5): 912-927. doi:10.12705/625.15

Wen, Jun, Xiong, Zhiqiang, Nie, Ze-Long, Mao, Likai, Zhu, Yabing, Kan, Xian-Zhao, Ickert-Bond, Stefanie, Gerrath, Jean, Zimmer, Elizabeth Anne and Fang, Xiao-Dong 2013. Transcriptome Sequences Resolve Deep Relationships of the Grape Family. PLoS ONE, 8(9): e74394 doi:10.1371/journal.pone.0074394

Zhang, Jing-Bo, Li, Rui-Qi, Xiang, Xiao-Guo, Manchester, Steven R., Lin, Li, Wang, Wei, Wen, Jun and Chen, Zhi-Duan 2013. Integrated Fossil and Molecular Data Reveal the Biogeographic Diversification of the Eastern Asian-Eastern North American Disjunct Hickory Genus (Carya Nutt.). PLoS ONE, 8(7): e70449 doi:10.1371/journal.pone.0070449

Zhou, Zhuo, Hong, Deyuan, Niu, Yang, Li, Guodong, Nie, Zelong, Wen, Jun and Sun, Hang 2013. Phylogenetic and biogeographic analyses of the Sino-Himalayan endemic genus Cyananthus (Campanulaceae) and implications for the evolution of its sexual system. Molecular phylogenetics and evolution, 68(3): 482-497. doi:10.1016/j.ympev.2013.04.027

Zhu, Wei-Dong, Nie, Ze-Long, Wen, Jun and Sun, Hang 2013. Molecular phylogeny and biogeography of Astilbe (Saxifragaceae) in Asia and eastern North America. Botanical Journal of the Linnean Society, 171(2): 377-394. doi:10.1111/j.1095-8339.2012.01318.x

Liu, Xiu-Qun, Ickert-Bond, Stefanie, Chen, Long-Qing and Wen, Jun 2012. Molecular phylogeny of Cissus L. of Vitaceae (the grape family) and evolution of its pantropical intercontinental disjunctions. Molecular phylogenetics and evolution, 66(1): 43-53. doi:10.1016/j.ympev.2012.09.003

Lu, Jin-Mei, Wen, Jun, Lutz, Sue M., Wang, Yi-Ping and Li, De-Zhu 2012. Phylogenetic relationships of Chinese Adiantum based on five plastid markers. Journal of Plant Research, 125(2): 237-249. doi:10.1007/s10265-011-0441-y

Lu, Limin, Wen, Jun and Chen, Zhiduan 2012. A combined morphological and molecular phylogenetic analysis of Parthenocissus (Vitaceae) and taxonomic implications. Botanical Journal of the Linnean Society, 168(1): 43-63. doi:10.1111/j.1095-8339.2011.01186.x

Mitchell, Anthony, Li, Rong, Brown, Joseph W., Schoenberger, Ines and Wen, Jun 2012. Ancient divergence and biogeography of Raukaua (Araliaceae) and close relatives in the southern hemisphere. Australian Systematic Botany, 25(6): 432-446. doi:10.1071/SB12020

Nie, Ze-Long, Funk, Vicki A., Sun, Hang, Deng, Tao, Meng, Ying and Wen, Jun 2012. Molecular phylogeny of Anaphalis (Asteraceae, Gnaphalieae) with biogeographic implications in the Northern Hemisphere. Journal of Plant Research, 126(1): 17-32. doi:10.1007/s10265-012-0506-6

Nie, Ze-Long, Sun, Hang, Manchester, Steven R., Meng, Ying, Luke, Quentin and Wen, Jun 2012. Evolution of the intercontinental disjunctions in six continents in the Ampelopsis clade of the grape family (Vitaceae). BMC Evolutionary Biology, 12(1): 1-17. doi:10.1186/1471-2148-12-17

Wen, Jun and Shi, Wenting 2012. Revision of the Maddenia clade of Prunus (Rosaceae). PhytoKeys, 11: 39-59. doi:10.3897/phytokeys.11.2825

Zhou, Zhuo, Wen, Jun, Li, Guodong and Sun, Hang 2012. Phylogenetic assessment and biogeographic analyses of tribe Peracarpeae (Campanulaceae). Plant Systematics and Evolution, 298(2): 323-336. doi:10.1007/s00606-011-0547-7

Zhu, Wei-Dong, Nie, Ze-Long, Wen, Jun and Sun, Hang 2012. Molecular phylogeny and biogeography of Astilbe (Saxifragaceae) in Asia and eastern North America. Botanical Journal of the Linnean Society, 171(2): 377-394. doi:10.1111/j.1095-8339.2012.01318.x

Zimmer, Elizabeth Anne and Wen, Jun 2012. Using nuclear gene data for plant phylogenetics: progress and prospects. Molecular phylogenetics and evolution, 65(2): 774-785. doi:10.1016/j.ympev.2012.07.015

Chen, P. T., Wen, Jun and Chen, L. Q. 2011. Spatial and temporal diversification of Tetrastigma Planch. (Vitaceae). Gardens' Bulletin Singapore, 63(1): 313-333.

Chen, Pingting, Chen, Longqing and Wen, Jun 2011. The first phylogenetic analysis of Tetrastigma (Miq.) Planch., the host of Rafflesiaceae. Taxon, 60: 499-512.

Hong, D. Y. and Wen, Jun 2011. Phrymaceae. In: Wu, C. Y., Hong, D. and Raven, Peter H., Flora of China. Beijing; St. Louis: Missouri Botanical Garden, pp.493-494.

Liu, Qing, Triplett, Jimmy K., Wen, Jun and Peterson, Paul M. 2011. Allotetraploid origin and divergence in Eleusine (Chloridoideae, Poaceae): evidence from low-copy nuclear gene phylogenies and a plastid gene chronogram. Annals of Botany, 108(7): 1287-1298. doi:10.1093/aob/mcr231

Lu, Jin-Mei, Li, De-Zhu, Lutz, Sue M., Soejima, Akiko, Yi, Tingshuang and Wen, Jun 2011. Biogeographic disjunction between eastern Asia and North America in the Adiantum pedatum complex (Pteridaceae). American Journal of Botany, 98(10): 1680-1963. doi:10.3732/ajb.1100125

Luebert, Federico, Brokamp, Grischa, Wen, Jun, Weigend, Maximilian and Hilger, Hartmut H. 2011. Phylogenetic relationships and morphological diversity in Neotropical Heliotropium (Heliotropiaceae). Taxon, 60: 663-680.

Ren, Hui, Lu, Li-Min, Soejima, Akiko, Luke, Quentin, Zhang, Dian-Xiang, Chen, Zhi-Duan and Wen, Jun 2011. Phylogenetic analysis of the grape family (Vitaceae) based on the noncoding plastid trnC-petN, trnH-psbA, and trnL-F sequences. Taxon, 60: 629-637.

Shui, Yu-Min, Wen, Jun and Chen, Wen-Hong 2011. Parashorea chinensis Wang Hsie and P. chinensis var. kwangsiensis Lin Chi: Two exceptional cases of names with a corporate authorship. Taxon, 60(4): 1165-1167.

Wen, Jun 2011. Systematics and Biogeography of Aralia L. (Araliaceae): Revision of Aralia Sects. Aralia, Humiles, Nanae, and Sciadodendron. Contributions from the United States National Herbarium, 57: 1-172.

Xie, Lei, Wen, Jun and Li, Liang-Qian 2011. Phylogenetic Analyses of Clematis (Ranunculaceae) Based on Sequences of Nuclear Ribosomal ITS and Three Plastid Regions. Systematic Botany, 36(4): 907-921. doi:10.1600/036364411X604921

Zhang, Jian-Wen, Nie, Ze-Long, Wen, Jun and Sun, Hang 2011. Molecular phylogeny and biogeography of three closely related genera, Soroseris, Stebbinsia, and Syncalathium (Asteraceae, Cichorieae), endemic to the Tibetan Plateau, SW China. Taxon, 60(1): 15-26.

Zuo, Y., Chen, Z., Kondo, K., Funamoto, T., Wen, Jun and Zhou, S. 2011. DNA barcoding of Panax species. Planta Medica, 72(2): 182-187. doi:10.1055/s-0030-1250166

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